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I have exactly the same problem. Spent last 2 days trying to figure out how to resolve this. Reinstalled my system about 10 times now. This definitely has to do with Bonjour service trying to start up but failing to set the policy correctly, so it just hangs. Unplugging network cable or disabling LAN card (or on-board adapter) doesn't make any difference. Still searching for a solution. Everything was working perfectly on GA-P35-DS4 Kalyway install since the beginning of this year. As soon as I upgraded to 10.5.5 things continued working fine until I copied new kexts when booting from another drive. My main machine is now well and truly broken because of this stupid mDNSResponder failure. No fix can be found anywhere yet (unfortunately).